THE HILL LAB
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Lab members

For a list of previous lab members:  Past Members
​

DR. MATTHEW HILL
Principal Investigator

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research interests
My research career has largely focused on the intersection between cannabinoids and the stress response. My lab has 3 primary research streams, all of which are independently funded, which focus on 1) stress and endocannabinoids; 2) cannabinoid regulation of feeding and metabolism; 3) the neurodevelopmental and behavioral impacts of cannabis exposure
education
I began working in the laboratory of Dr. Boris Gorzalka as a 2nd year undergraduate in 1999. My initial projects were at looking at the behavioural effects of the hormones corticosterone and melatonin on stress coping behaviour. I continued graduate work in the Gorzalka lab where I transitioned into exploring the endocannabinoid system, how it interacts with stress hormones and how these changes contribute to alterations in affective and cognitive behaviour. The focus of my thesis was examining the impacts of chronic stress and antidepressant treatments on the endocannabinoid system. I also spent a large portion of my graduate training in two other labs, those of Dr. Cecilia Hillard and Dr. Victor Viau, where I honed skills in the neuropharmacology of cannabinoids and the psychoneuroendocrinology of stress, respectively. In 2008, I finished my PhD and left UBC to join the laboratory of Dr. Bruce McEwen at Rockefeller University in New York City. I finished my postdoctoral fellowship and started my own lab at the University of Calgary in the summer of 2011.
hobbies
​Personally, I love to cook and travel and am an avid dog fan, especially those of the bulldog breed. I like to ski and hike in the rocky mountains nearby
favorite paper
​Favourite paper: I am going to select 3 papers, as there were 3 key papers that got the entire field of stress, affective behavior and endocannabinoids rolling:
1)    Marsicano G, Wotjak CT, Azad SC, Bisogno T, Rammes G, Cascio MG, Hermann H, Tang J, Hofmann C, Zieglgänsberger W, Di Marzo V, Lutz B.  (2002) The Endogenous Cannabinoid System Controls Extinction of Aversive Memories. Nature 418(6897): 530-4.
2)    Di S, Malcher-Lopes R, Halmos KC, Tasker JG. (2003). Nongenomic glucocorticoid inhibition via endocannabinoid release in the hypothalamus: a fast feedback mechanism. J Neurosci. 23(12): 4850-7.
3)    Patel S, Roelke CT, Rademacher DJ, Cullinan WE, Hillard CJ. (2004). Endocannabinoid signaling negatively modulates stress-induced activation of the hypothalamic-pituitary-adrenal axis. Endocrinology 145(12): 5431-8.
publications
SELECT PUBLICATIONS
Stress-induced Modulation of Endocannabinoid Signaling Leads to Delayed Strengthening of Synaptic Connectivity in the Amygdala.
Yasmin F, Colangeli R, Morena M, Filipski S, Pittman QJ, van der Stelt M, Hillard CJ, Teskey GC, McEwen BS, Hill MN*, Chattarji S*. (2020)
Proceeding of the National Academy of Sciences USA 117(1), 650-655.
* = co-senior authorship
​LINK

Neurobiological Interactions Between Stress and the Endocannabinoid System
Morena M, Patel S, Bains JS, Hill MN (2016)
Neuropsychopharmacology 41(1), 80-102
​LINK
 
Corticotropin Releasing Hormone Drives Anandamide Hydrolysis in the Amygdala to Promote Anxiety.
Gray JM, Vecchiarelli HA, Morena M, Lee TT, Hermanson D, Kim AB, McLaughlin RJ, Hassan K, Kuhne C, Wotjak CT, Deussing JM, Patel S, Hill MN (2015)
Journal of Neuroscience 35(9), 3879-3892
​LINK

 
Full list of publications​
awards
  • 2020: Gill Center Transformative Research Award – Gill Center for Biomolecular Science at Indiana University
  • 2018-2019:  Hungarian Academy of Sciences Visiting Professorship – Institute of Experimental Medicine
  • 2018: International Cannabinoid Research Society Young Investigator of the Year Award
  • 2017: Elected into the Royal Society of Canada in the College of New Scholars, Artists and Scientists
  • 2017: International Association for Cannabinoid Medicines Young Research Award
  • 2016: Killam Emerging Research Leader Award
  • 2016: Canadian Association for Neuroscience Young Investigator of the Year Award
  • 2014: Top 40 Under 40, Calgary, Avenue Magazine
  • 2012-2022: Tier II Canada Research Chair in the Neurobiology of Stress
  • 2010: Best Oral Presentation (Post-Doctoral), 20th Annual Symposium of the International Cannabinoid Research Society
  • 2010: Travel Award, 49th Annual Meeting of the American College of Neuropsychopharmacology
Contact: mnhill@ucalgary.ca

MIN QIAO
Lab Manager & Technician (2015 - present)

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role
I am the lab manager and research technician in the lab. I joined the lab in 2015 and has been responsible for lab safety and supplies procurement. I provide routine technical assistance in the lab and have many years of expertise in immunohistology, Westen Blot and rodent surgeries.
education
​​I completed my Medical degree from Southwest Medical University, Peking Union Medical College and Chinese Academy of Medical Sciences (1988-1996). I moved to Canada and received my Msc in Neurophysiology from University of Manitoba in 1999 under the supervision of Dr. Ursula Tuor.  After graduation, I worked for 10 years at Institute for Biodiagnostics, National Research Council Canada in the research of magnetic resonance imaging on rodent models of stroke before working at the University of Calgary.
hobbies
​In my free time, I like to cook and play sports such as ping-pong, badminton, and soccer with my family. ​
publications
​Full list of publications
Contact: ​mqiao@ucalgary.ca

DR. GEORGIA BALSEVICH
Postdoctoral Fellow (2016 - present)

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research interests
​I am interested in understanding the crosstalk between energy balance and stress regulation by identifying underlying shared mechanisms. In the Hill lab, I am specifically investigating glucocorticoid-endocannabinoid signaling mechanisms that regulate whole body energy metabolism by using a combination of molecular, genetic, and pharmacological approaches.
education
​I completed my BSc in Biochemistry at the University of Saskatchewan before receiving my MSc in Neuroscience from the University of British Columbia. I then moved to Munich, Germany for my PhD in order to work with Dr. Mathias Schmidt at the Max Planck Institute of Psychiatry where I investigated molecular mechanisms contributing to disease susceptibility. Indeed my current research interests have been shaped by my past research experience, which span molecular, cellular, and systems-level examination of stress-related pathophysiologies.
hobbies
​​In my free time, I love to be on the move, be it in the city running and cycling, or in the mountains hiking and skiing.
favorite paper
​​I cannot pick just one because too many papers have inspired my research over the years, which is continuously evolving. However, a favourite pick from the last two years is “Activation of AMPK-Regulated CRH Neurons in the PVH is Sufficient and Necessary to Induce Dietary Preference for Carbohydrate over Fat” (Okamoto et al, 2018; PubMed link). I like this paper because the authors pinpoint a cell-specific molecular mechanism to explain a highly complex behaviour in order to help us understand why food preference shifts in response to the environmental context (i.e. metabolic states and stress).
publications
​SELECT PUBLICATIONS
A Role for Fatty Acid Amide Hydrolase (FAAH) in the Leptin-mediated Effects on Feeding and Metabolism
Balsevich G, Sticht MA, Bowles NP, Singh A, Lee TT, Li Z, Chelikani P, Lee FS, Borgland SL, Hillard CJ, McEwen BS, Hill MN (2018)
Proceedings of the National Academy of Sciences USA 115(29), 7605-7610.
​LINK

Stress-responsive  FKBP51 regulates AKT2-AS160 signaling and metabolic function
Balsevich G, Häusl AS, Meyer CW, Karamihalev S, Feng X, Pöhlmann ML, Dournes C, Uribe-Marino A, Santarelli S, Labermaier C, Hafner K, Mao T, Breitsamer M, Theodoropoulou M, Namendorf C, Uhr M, Paez-Pereda M, Winter G, Hausch F, Chen A, Tschöp MH, Rein T, Gassen NC, Schmidt MV (2017)
Nature Communications, 8(1), 1725
​LINK

Endocannabinoids: Effectors of Glucocorticoid Signaling
Balsevich G, Petrie GN, Hill MN (2017)
Frontiers in Neuroendocrinology 47, 86-108
​LINK


Full list of publications
awards
​- 2019: Awarded Alberta Innovates Postgraduate Fellowship
​- 2018: Awarded Banting Postdoctoral Fellowship, most prestigious and competitive fellowship of those offered by the Canadian Federal Government
- 2018: Awarded Canadian Institutes of Health Research (CIHR) Postdoctoral Fellowship, ranked 1st out of 1071 applications submitted, declined
- 2017 – 2018: Elected Director of Events, The Postdoctoral Association, University of Calgary, Calgary
- 2017: Awarded Postdoctoral Presentation Award, International Cannabinoid Research Society
- 2016: Awarded Killam Postdoctoral Fellowship
- 2015 – 2016: Elected PhD representative for the Max Planck Society, Munich, Germany
- 2014: Awarded Young Investigator Award at the 27th European College of Neuropsychopharmacology (ECNP) Congress Berlin
- 2012: Selected member of the International Max Planck Research School-LS, Munich, Germany
- 2011: Awarded NSERC CGSM Scholarship
Contact: georgia.balsevich@ucalgary.ca

DR. CATHERINE HUME
Postdoctoral Fellow (2019 - present)

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research interests
​I am interested in the neuronal control of homeostatic and hedonic feeding behaviours and the modulation of these circuits by environmental factors that alter behaviour. I am currently investigating how cannabis stimulates food intake (a phenomenon commonly known as ‘the munchies’) with the aim of deciphering the neuronal mechanisms that underlie these cannabis-induced motivated feeding behaviours. This research involves the use of specialized cannabis vapor administration chambers combined with various behavioural paradigms (including operant conditioning), as well as pharmacological and circuit-based manipulations.​
education
After completing my BSc in pharmacology, I remained at the University of Edinburgh (UK) to carry out my PhD in the Centre for Integrative Physiology with Professor Gareth Leng and Dr John Menzies. I investigated how the homeostatic energy balance system compensates for the hedonic consumption of palatable, high-sugar food to maintain body weight.
Following a short postdoc in the lab of Professor Mike Ludwig (Centre of Discovery Brain Sciences, University of Edinburgh (UK)) investigating how retinal vasopressin regulates hypothalamic circuits controlling circadian rhythms, I moved to Dr. Matthew Hill’s lab at the University of Calgary to continue pursuing feeding behaviour-based research.
hobbies
I like to cook, pet dogs, watch true crime documentaries and drink margaritas :) ​​
favorite paper
​​‘AGRP neurons are sufficient to orchestrate feeding behavior rapidly and without training’ (Aponte et al 2011 Nat Neurosci; PubMed link). This paper has a special place in my heart as after reading it, I decided to carry out a PhD in feeding behaviour research. It is an exceptional study showing for the first time that feeding behaviours can be driven by the specific activation of a single hypothalamic circuit.
publications
SELECT PUBLICATIONS
Effects of optogenetic stimulation of vasopressinergic retinal afferents on suprachiasmatic neurones
Hume C, Allchorne A, Grinevich V, Leng G, Ludwig M (2019)
Journal of Neuroendocrinology 31(12):e12806
LINK

High-Sugar, but Not High-Fat, Food Activates Supraoptic Nucleus Neurons in the Male Rat
Hume C, Sabatier N, Menzies J
Endocrinology 158(7):2200-2211
​LINK

Homeostatic responses to palatable food consumption in satiated rats
Hume C, Jachs B, Menzies J
Obesity (Silver Spring) 24(10):2126-32

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Maternal-fetal transmission of delta-9-tetrahydrocannabinol (THC) and its metabolites following inhalation and injection exposure during pregnancy in rats
Baglot SL, VanRyzin JW, Marquardt AE, Aukema RJ, Petrie GN, Hume C, Reinl EL, Bieber JB, McLaughlin RJ, McCarthy MM, Hill MN. (2021)
Journal of Neuroscience Research, published ahead of print
​LINK


Full list of publications​
awards
  • Sept 2019: Awarded Eyes High Postdoctoral Scholarship
  • April 2019: Invited speaker for British Neuroscience Association Festival of Neuroscience in Dublin, Ireland
  • Oct 2018: ‘Named Postdoctoral Researcher’ on large BBSRC grant (BB/S000224/1) awarded to the University of Edinburgh, UK 
  • Sept 2017: Selected as committee member for George Square Postdoc Society at the University of Edinburgh, UK
  • Feb 2016: Received ‘Associate Fellow’ status by the Higher Education Academy UK for developing my teaching repertoire at the University of Edinburgh, UK
  • July 2015: Selected to represent the University of Edinburgh at the League of European Research Universities Doctoral Summer School hosted by the University of Oxford, UK
  • Jan 2014: Elected as ‘Early Stage Career Representative’ for the large-scale European Commission-funded project ‘Nudge-it’
  • July 2012: Awarded Wellcome Trust Biomedical Vacation Scholarship
Contact: ​catherine.hume1@ucalgary.ca

​ROBERT AUKEMA
PhD Candidate (2016 - present)

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research interests
I am interested in understanding basolateral amygdala circuitry during exposure to stress. Broadly, I am investigating the anatomy (topographical distribution) and the functionality of projection neuron populations in the basolateral amygdala to understand the contribution of each population to physiological and behavioral responses to stress challenge. To answer these questions, I use retrograde tracing, immunohistochemistry, Gi DREADDs, optogenetics, fiber photometry, and behavioral tests. 
education
​I received my undergraduate degree (B.Sc. 2012-2016) at Carleton University in Ottawa, ON, where I completed an Honor’s Thesis in Dr. Alfonso Abizaid’s lab – a place that first truly fuelled my interest in neuroscience. ​
hobbies
Outside of the lab, I am a whole-hearted fan of the mountains, my 2006 Hyundai Sante-Fe, the squash court, and the Toronto Maple Leafs.​
favorite paper
​“Organization of valence-encoding and projection-defined neurons in the basolateral amygdala” (Beyeler et al, 2018; PubMed link). A beautiful and detailed anatomical description of three projection neuron populations in the amygdala, and how they interact to encode valence of stimuli.
publications
SELECT PUBLICATIONS
Endocannabinoids, cannabinoids and the regulation of anxiety

Petrie GN, Nastase AS, Aukema RJ, Hill MN (2021)
Neuropharmacology 195:108626
​LINK

​Up-regulation of Anandamide Hydrolysis in the Basolateral Complex of the Amygdala Reduces Fear Memory Expression and Indices of Stress and Anxiety.
Morena M, Aukema RJ, Leitl K, Rashid A, Vecchiarelli HA, Josselyn SA, Hill MN (2019).
Journal of Neuroscience 39(7), 1275-1292.
​LINK


Protective Effects of Elevated Anandamide on Stress and Fear-related Behaviors: Translational Evidence from Humans and Mice
Mayo LM, Asratian A, Linde J, Holm L, Natt D, Augier G, Stensson N, Vecchiarelli HA, Balsevich G, Aukema RJ, Ghafouri B, Spagnolo PA, Lee FS, Hill MN, Heilig M (2020)
Molecular Psychiatry 25(5):993-1005
​LINK
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​Full list of publications
Awards
I am extremely grateful to have received funding support from:
  • Mathison Centre Graduate Scholarship (2018-present), Mathison Centre for Mental Health & Education
  • CSM Graduate Student Scholarship (2017-2018), Cumming School of Medicine
Contact: robert.aukema1@ucalgary.ca​       /        ​Twitter: ​@robertaukema

​GAVIN PETRIE
PhD Candidate (2016 - present)

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research interests
​My research investigates how the endocannabinoid system gates the stress response.  Specifically, I am investigating how both tonic and phasic endocannabinoid signalling regulate stress-responsivity by measuring HPA axis activation and self-directed grooming behaviors.  I am harnessing a wide range of techniques to answer these questions: pharmacology, optogenetics, fiber photometry and immunohistochemistry.
education
​​I earned my Undergraduate degree in psychology at the University of Guelph (B.A. 2016).  My passion for cannabinoids began while completing an honors thesis in Linda Parker’s lab on how the endogenous cannabinoid, Oleoyl Glycine, can be used to treat morphine withdrawal.
hobbies
​​My recreational passions include mountain biking, skiing and having a pint with my buddies.
favorite paper
​​Favorite paper: “Nongenomic Glucocorticoid Inhibition via Endocannabinoid Release in the Hypothalamus: A Fast Feedback Mechanism” (Di et al., 2003; PubMed link).  This is the first paper to suggest a role for endocannabinoids in the stress response by showing the necessity of cannabinoid signalling during glucocorticoid negative feedback in the PVN.
publications
SELECT PUBLICATIONS​
Endocannabinoids, cannabinoids and the regulation of anxiety
Petrie GN, Nastase AS, Aukema RJ, Hill MN (2021)
Neuropharmacology 195:108626
​LINK

Oleoyl glycine: interference with the aversive effects of acute naloxone-precipitated MWD, but not morphine reward, in male Sprague-Dawley rat
Petrie GN, Wills KL, Piscitelli F, Smoum R, Limebeer CL, Rock EM, Humphrey AE, Sheppard-Perkins M, Lichtman AH, Mechoulam R, Di Marzo V, Parker LA
Psychopharmacology (Berlin), 236(9):2623-2633
​LINK

​Discovery of a NAPE-PLD inhibitor that modulates emotional behavior in mice.
Mock  ED, Mustafa M, Gunduz-Cinar O, Cinar R, Petrie GN, Kantae V, Di X, Ogasawara D, Varga Z, Paloczi J, Miliano C, Donvito G, van Esbroeck ACM, van der Gracht AMF, Kotsogianni L, Park JK, Martella A, van der Wel T, Soethoudt M, Jiang M, Wendel TJ, Janssen APA, Bakker AT, Donovan CM, Castillo LI, Florea BI, Wat J, van den Hurk H, Wittwer M, Grether U, Holmes A, van Boeckel CAA, Hankemeier T, Cravatt BF, Buczynski MW, Hill MN, Pacher P, Lichtman AH, van der Stelt  M (2020)
Nature Chemical Biology 16(6):667-675
​LINK

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Full list of publications
awards
​Awards
  • Third Place Poster, HBI Research Day (2019); Hotchkiss Brain Institute
  • Billy Martin Award (2018); International Cannabinoid Research Society
  • Honorable Mention, Graduate Newcomer Excellence Award (2017); University of Calgary Graduate Student Association ​
Funding
  • Doctoral Scholarship (2019-present), BranchOut Neurological Foundation
  • CSM Graduate Student Scholarship (2017-2019), Cumming School of Medicine
  • Canadian Graduate Scholarship for Masters (2017-2018), CIHR
  • Queen Elizabeth II Scholarship (2016), Tri-Council Canada
Contact: ​gavin.petrie@ucalgary.ca

SAMANTHA BAGLOT
PhD Candidate (2018 - present)

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RESEARCH INTERESTS
​Overall, my research investigates the endocrine, immune, and behavioural effects of developmental exposure to cannabis. Utilizing cannabis vapour administration chambers my research specifically examines the effects of either prenatal or adolescent exposure to cannabis on stress and immune system development and functioning, as well as social behaviour.
EDUCATION
​​I completed my BA in Psychology (2011-2015) and MSc in Neuroscience (2015-2018) from the University of British Columbia. During my MSc degree I was co-supervised by Drs. Joanne Weinberg and Liisa Galea and I investigated a possible role for oxytocin in attenuating the effects of prenatal alcohol exposure on adult stress functioning, hippocampal neurogenesis, and anxiety-like behaviour.
HOBBIES
​In my free time, I love to cycle (both outdoors or in a spin class), hike, practice yoga, and ski. I am a second-degree black belt in Meibukan Go-Ju Ryu Karate and a first-degree in Kobudo (Weapons).
FAVORITE PAPER
​​Favorite paper: “Microglial Phagocytosis of Newborn Cells Is Induced by Endocannabinoids and Sculpts Sex Differences in Juvenile Rat Social Play” (VanRyzin et al., 2019; PubMed link). This paper highlights the developmental and sex-specific role of the endocannabinoid system in promoting the immune system to shape the circuity underlying social play behaviour - and is the first paper that got me interested in how prenatal cannabis exposure would alter immune system development and social behaviour functioning.
PUBLICATIONS
SELECT PUBLICATIONS​
Maternal-fetal transmission of delta-9-tetrahydrocannabinol (THC) and its metabolites following inhalation and injection exposure during pregnancy in rats
Baglot SL, VanRyzin JW, Marquardt AE, Aukema RJ, Petrie GN, Hume C, Reinl EL, Bieber JB, McLaughlin RJ, McCarthy MM, Hill MN. (2021)
Journal of Neuroscience Research, published ahead of print
​LINK


Cannabis vapor self-administration elicits sex- and dose-specific alterations in stress reactivity in rats
Glodosky NC, Cuttler C, Freels TG, Wright HR, Rojas MJ, Baglot SL, Hill MN, McLaughlin RJ (2021).
Neurobiology of Stress 13:100260
​LINK

Effects of prenatal alcohol exposure on social competence: Asymmetry in play partner preference among heterogeneous triads of male and female rats
Holman PJ, Baglot SL, Morgan E, Weinberg J (2019)
Developmental Psychobiology 61(4):513-524
​LINK


Full list of publications
Awards
Awards
  • Graduate Citizenship Award  (2019); University of Calgary Graduate Student Association
  • Best Trainee Talk at Perinatal Origins of Neuropsychiatric Disorders (POND) Conference (2019)
  • Young Investigator Trainee Program Award (2019); Perinatal Origins of Neuropsychiatric Disorders (POND) Conference​
Funding
  • CIHR Vanier Doctoral Award (2018-present), Canadian Institute of Health Research
  • Graduate Recruitment Scholarship in Mental Health (Declined 2018), Mathison Centre 
  • Aboriginal Graduate Fellowship and Tuition Award (2015-2017), University of British Columbia
  • NSERC Canada Graduate Scholarship Master's Award (2015-2016), Natural Sciences & Engineering Research Council
  • Queen Elizabeth II Graduate (Master's) Scholarship (Declined 2015), University of Calgary​
Contact: ​samantha.baglot@ucalgary.ca       /       Twitter: @s_baglot

​ANDREI SABIN NASTASE
PhD/MD Candidate (2018 - present)

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RESEARCH INTERESTS
​My research focuses on how the endocannabinoid system acts to regulate valence processing and coordinate behavioral invigoration during stress and homeostatic need. My approach is rooted in an ethological framework; by evaluating behavioral repertoires in a semi-naturalistic approach/avoidance task, I seek to answer my question in an evolutionarily relevant way.
EDUCATION
​I earned my Bachelor of Science (BSc) Honours Degree in Neuroscience at the University of Calgary. My first exposure to research was at the Alberta Children’s Hospital under the supervision of Dr. Daniele Pacaud, where I studied the effectiveness of Corneal Confocal Microscopy in predicting and detecting pediatric diabetic neuropathy. My passion for research blossomed during my honours thesis under the supervision of Dr. Matt Hill and Dr. Maria Morena, with whom I investigated the influence of the endocannabinoid system in regulating fear memory dynamics. I am currently enrolled in the Leader's in Medicine Program at the University of Calgary.
HOBBIES
I love playing frisbee, riding my bike, reading classics, and engaging in as much banter as I can.​
FAVORITE PAPER
​Favorite paper:  “Amygdala Signaling during Foraging in a Hazardous Environment” (Amir et al., 2015). Great use of an ethological model to explore the behavioral correlates of neuronal activity in a sub-section of basolateral amygdala neurons.
Publications
Sex dependent effects of endocannabinoid modulation of fear extinction in rats.
Morena M, Nastase AS, Santori A, Cravatt BF, Shansky RM, Hill MN. (2021)
British Journal of Pharmacology 178(4), 983-996

​LINK

Endocannabinoids, cannabinoids and the regulation of anxiety
Petrie GN, Nastase AS, Aukema RJ, Hill MN (2021)
Neuropharmacology 195:108626
​LINK


Amygdalar endocannabinoids are affected by predator odor stress in a sex-specific manner and modulate acoustic startle reactivity in female rats
Albrechet-Souza L, Nastase AS, Hill MN, Gilpin NW (2021)
Neurobiology of Stress 15:100387
LINK


​Full list of publications
AWARDS
​Canada Graduate Scholarships-Master’s Program (Canadian Institutes of Health
Research – Fredrick Banting and Charles Best Canada Graduate Scholarships) – 2019-2020
 
Cumming School of Medicine Graduate Scholarship – 2019-2021
 
Mathison Centre Graduate Recruitment Scholarship in Mental Health - 2019-2021
 
Hotchkiss Brain Institute Graduate Recruitment Scholarship in Neuroscience (awarded and declined)
 
Branch Out Neurological Foundation Undergraduate Studentship – 2017
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Markin Undergraduate Student Research Program in Health and Wellness Award (awarded and declined)
​Contact: andrei.nastase@ucalgary.ca
Research funded by Brain Canada, BONF, CFI, CIHR, NIH and NSERC. Many thanks to the University of Calgary Cumming School of Medicine, Hotchkiss Brain Institute, Mathison Centre, and Alberta Children's Hospital for their support.
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